Timothy D. Schowalter




Insect Ecology and Forest Entomology.


EMAIL: tschowalter@agcenter.lsu.edu


DEGREES:     B.A. 1974 Wichita State University

                      M.S. 1976 New Mexico State University

                      Ph.D. 1979 University of Georgia


Dr. Schowalter is Professor of Entomology at Louisiana State University (2003-present) and served as Department Head 2003-2015.  He was a professor at Oregon State University for 22 yrs, (1981-2003), serving as Interim Head (2001-2003) before moving to LSU.  He has published four books, more than 70 peer-reviewed journal papers, 3 invited reviews, and a large number of additional book chapters, symposia proceedings and other publications.  He was Program Director for Ecosystem Studies at the National Science Foundation (1992-1993) and has served as Vice President for Public Affairs, Ecological Society of America (1999-2002) and Governing Board member for the Entomological Society of America (2013-2016).  He was named a Fellow of the Ecological Society of America in 2012.


Current Courses:

ENTM 4018 Forest Insects and Disease

ENTM 4040 Insect Ecology


Current Research Interest:


I am continuing long-term studies of canopy arthropod responses to hurricanes and other disturbances in forest ecosystems.  I now have twenty five years of data for post-Hugo (1989) and post-Georges (1998) recovery of canopy arthropod communities in tropical rainforests at the Luquillo Experimental Forest Long Term Ecological Research (LTER) Site in Puerto Rico.  I am collaborating with colleagues at Oregon State University, National Taiwan University, and the Taiwan Forest Research Institute on comparison of the data from Puerto Rico with comparable data following typhoon disturbances in sub-tropical forests in Taiwan.  Following Hurricanes Katrina (2005) and Gustav (2008), I initiated studies in lowland hardwood forests in southeastern Louisiana.  The combination of these databases from different forest types will improve our prediction of canopy arthropod responses to canopy-opening disturbances.  I currently am addressing the ecology and management of the emerald ash borer, which recently invaded Louisiana.



Research Databases:








Publications (since 2000):


Peer-Reviewed Journals (since 2000)


Rambo, T., T. Schowalter and M. North. 2014. Canopy arthropod responses to thinning and burning treatments in old-growth mixed-conifer forest in the Sierra Nevada, California. Forest Ecology and Management 326: 91-100.


Schowalter, T.D., M.R. Willig and S.J. Pressley. 2014. Canopy arthropod responses to experimental canopy opening and debris deposition in a tropical rainforest subject to hurricanes. Forest Ecology and Management 332: 93-102.


Schowalter, T.D. 2014. Ecology and management of bark beetles. Formosan Entomologist 33: 167-187.


Schowalter, T.D. 2012. Ecology and management of bark beetles (Coleoptera: Curculionidae: Scolytinae) in southern pine forests. Journal of Integrated Pest Management 3(2): A1-A7.


Schowalter, T.D. 2012. Insect herbivore effects on forest ecosystem services. Journal of Sustainable Forestry 31: 518-536.


Schowalter, T.D., S.J. Fonte, J.Geagan and J. Wang. 2011. Effects of manipulated herbivore inputs on nutrient flux and decomposition in a tropical rainforest in Puerto Rico. Oecologia 167:1141–1149.


Schowalter, T.D. 2008. INSECT herbivore responses to management practices in conifer forests in North America. Journal of SUSTAINABLE FORESTRY 26: 204-222.


Fonte, S.J. and T.D. Schowalter. 2005. The influence of a neotropical herbivore (Lamponius portoricensis) on nutrient cycling and soil processes. Oecologia 146: 423-431.


Schowalter, T.D. and Y.L. Zhang. 2005. Canopy arthropod assemblages in four overstory and three understory plant species in mixed-conifer old-growth forest in California. Forest Science 51: 233-242.


Schowalter, T.D., Y.L. Zhang and R.A. Progar. 2005. Canopy arthropod response to density and distribution of green trees retained after partial harvest. Ecological Applications 15: 1594-1603.


Fonte, S.J. and T.D. Schowalter. 2004. Decomposition of greenfall vs. senescent foliage in relation to herbivory and disturbance in a forest ecosystem. Biotropica 36: 374-382.


Schowalter, T.D., Y.L. Zhang and J.J. Rykken. 2003. Litter invertebrate responses to variable density thinning in western Washington forest. Ecological Applications 13: 1204-1211.


Progar, R.A. and T.D. Schowalter. 2002. Canopy arthropod assemblages along a precipitation and latitudinal gradient among Douglas-fir Pseudotsuga menziesii forests in the Pacific Northwest of the United States. Ecography 25: 129-138.


Schowalter, T.D. and J.J. Morrell.  2002.  Nutritional quality of Douglas-fir wood: effect of vertical and horizontal position on nutrient levels.  Wood and Fiber Science 34: 158-164.


Schowalter, T.D. and J. Withgott. 2001. Rethinking insects: what would an ecosystem approach look like?  Conservation Biology in Practice 2: 10-16.


Smith, J.P. and T.D. Schowalter. 2001. Aphid-induced reduction in shoot and root growth of Douglas-fir seedlings. Ecological Entomology 26: 411-416.


Progar, R., T.D. Schowalter, J.J. Morrell and C.M. Freitag. 2000. Respiration from coarse woody debris as affected by moisture and saprotroph functional diversity in western Oregon. Oecologia 124: 426-431.


Reviews (since 2000)


Lowman, M.D. and T.D. Schowalter. 2012. Tansley Review: Plant science in forest canopies – the first forty years of advances and challenges (1980 – 2010). New Phytologist 194: 12-27.


Schowalter, T.D. 2012. Insect responses to major landscape-level disturbance. Annual Review of Entomology 57: 1-20.


Books and Book Chapters (since 2000)


Reynolds, B.C., T.D. Schowalter and D.A. Crossley, Jr. 2014. Watershed clearcutting and canopy arthropods. pp. 146-155 In W.T. Swank and J.R. Webster, eds., Long-term Response of a Forest Watershed Ecosystem. Oxford University Press, New York.


Schowalter, T.D. 2013. Insects and Sustainability of Ecosystem Services. CRC Press/Taylor and Francis Group, Boca Raton, FL.


Lowman, M.D., T.D. Schowalter and J.F. Franklin. 2012. Methods in Forest Canopy Research. University of California Press, Berkeley.


Schowalter, T.D. 2012. Insect outbreak effects on ecosystem services, pp. 246-265 In P. Barbosa, D.K. Letourneau and A.A. Agrawal, eds., Insect Outbreaks Revisited. Wiley/Blackwell.


Schowalter, T.D. 2011. Insect Ecology: an Ecosystem Approach, 3rd Ed. Elsevier/Academic, San Diego, CA.


Schowalter, T.D. 2007. Ecosystems: concepts, analyses and practical implications, Pp. 411-430, In: M. Kogan and P. Jepson, eds., Perspectives in Ecological Theory and Integrated Pest Management. Cambridge University Press, Cambridge, U.K.


Schowalter, T.D. 2006. Insect Ecology: an Ecosystem Approach, 2 nd Ed. Elsevier/Academic, San Diego, CA.


Fonte, S.J. and T.D. Schowalter. 2004.  Decomposition in forest canopies. pp. 413-422, In:  M.D. Lowman and B. Rinker, eds., Forest Canopies. Elsevier, Amsterdam.


Schowalter, T.D. and L.M. Ganio. 2003.  Diel, seasonal and disturbance-induced variation in invertebrate assemblages.  pp. 315-328, In: Y. Basset, V. Navotny, S. Miller and R. Kitching, eds., Arthropods of Tropical Forests. Cambridge University Press, Cambridge, UK.


Christensen, N.L., Jr., S.V. Gregory, P.R. Hagenstein, T.A. Heberlein, J.C. Hendee, J.T. Olson, J.M. Peek, D.A. Perry, T.D. Schowalter, K. Sullivan, G.D. Tilman and K.A. Vogt. 2000. Environmental Issues in Pacific Northwest Forest Management. National Academy Press, Washington, D.C.